The USS Titan-A is an Epic Grade 4 Explorer, a Neo-Constitution design inspired by older Constitution-class ships. Its overpowered impulse engines give it the largest sub-light power-to-geometry ratio in Starfleet. To build it you need a Shipyard at level 37 and Tolerated standing with the Ex-Borg Faction. At Tolerated, you will see a bundle with the blueprints.
The Titan-A is the first ship with two active abilities that boost your ships and your Alliance's ships. They help in hostile hunting, PvP, Territory Capture, and Armadas. Both abilities also counter the Mantis's Venomous Puncture. Fortification applies a Fortified state to every ship you have in the same system, plus some Alliance ships. The state gives 25% Critical Damage and activates some of the Titan research buffs the receiving player has unlocked. Max Fortification gives 25% Critical Damage and 250% Damage, and activates all unlocked Titan research buffs. Both abilities cost Radioactive Titanite. You get it from the Gifts tab once the ship is built, and the amount grows as the ship tiers up. Both have a duration and a cooldown. Alliance ships are picked by highest Attack stat first. A ship already in one of the states is skipped. If one of your own ships is already in a state, its duration is refreshed.
Tiering up the Titan-A shortens the cooldown and lengthens the duration of its abilities. Titan-A research nodes improve them further and can add new effects. Unlocked nodes apply only to your own ships, not to other players, the same rule as for the Cerritos and Defiant. Veridium Isotope unlocks the research nodes. You get it from the ability daily goals and from the Offers tab. The ship upgrades with USS Titan-A Parts from the Fortification Daily Goal and the Offers tab. Based on tier, some ships can benefit without counting against the normal ship limit. Both abilities only affect undocked ships.